Proper Focus . Answers typically involve functional, environmental, and, increasingly, developmental factors that influence the dynamics of natural selection and genetic drift. But what about evolutionary developmental biology? What is its characteristic question type? If we are correct that a genuinely distinctive scientific discipline must have a distinctive question to ask, then in order for an inquiry to count as part of evo‐devo, it must pursue an inquiry that is not wholly in the compass of evolutionary or developmental biology. As noted in ‘Introduction’, we suggest that the distinguishing questions for evolutionary developmental biology concern evolvability; these are questions about how variation is generated. Before pushing this idea any further, however, it is worth exploring our suggestion that a scientific discipline is distinguished by a characteristic central question. The idea of a central question The idea that a new scientific discipline must have a novel sort of question to ask bears much of the load of our argument. It allows us to make the uncomfortable suggestion that not every party waving the evo‐devo flag produces genuine evo‐devo work, .